Post by Barry the Baryonyx on Jun 26, 2007 0:33:33 GMT -5
Tyrannosaurus rex
Tyrannosaurus meaning 'tyrant lizard') is a genus of tyrannosaurid theropod dinosaur. The species Tyrannosaurus rex, commonly abbreviated to T. rex, is one of the dinosaurs most often featured in popular culture around the world. It hails from what is now western North America. Some scientists consider the slightly older Tarbosaurus bataar from Asia to represent a second species of Tyrannosaurus, while others maintain Tarbosaurus as a separate genus.
Like other tyrannosaurids, Tyrannosaurus was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to the large and powerful hindlimbs, Tyrannosaurus forelimbs were small and retained only two digits. Although other theropods rivaled or exceeded T. rex in size, it was the largest known tyrannosaurid and one of the largest known land predators, measuring over 12 metres (40 feet) in length and weighing as much as an elephant.
Fossils of some T. rex have been found in North American rock formations dating to the very end of the Cretaceous Period (late Maastrichtian stage, 65 million years ago); it was among the last dinosaurs to exist prior to the Cretaceous-Tertiary extinction event. More than 30 specimens of T. rex have now been identified, some nearly complete, which has allowed significant research into many aspects of its biology, including its life history and biomechanics. The feeding habits and potential speed of T. rex remain controversial.
Description
Tyrannosaurus rex was one of the largest land carnivores of all time, about 12 to 13 meters (40 to 43.3 feet) long, and 4.5-5 m (14-16 ft) tall, when fully-grown.[1] Mass estimates have varied widely over the years, from more than 7,200 kilograms (8 short tons), to less than 4,500 kg (5 tons), with most modern estimates ranging between 5,400 and 6,800 kg (between 6 and 7.5 tons).
The largest known T. rex skulls measure up to 1.5 m (5 ft) in length. Compared to other theropods, the skull was heavily modified. The skull was extremely wide posteriorly, with a narrow snout, allowing some degree of binocular vision. Some of the bones, such as the nasals, were fused, preventing movement between them. Large fenestrae (openings) in the skull reduced weight and provided areas for muscle attachment. The bones themselves were massive, as were the serrated teeth which, rather than being bladelike, were oval in cross-section. Like other tyrannosaurids, T. rex displayed marked heterodonty, with the premaxillary teeth at the front of the upper jaw closely-packed and D-shaped in cross-section. Large bite marks found on bones of other dinosaurs indicate that these teeth could penetrate solid bone. T. rex had the greatest bite force of any animal. Worn or broken teeth are often found, but unlike those of mammals, tyrannosaurid teeth were continually replaced throughout the life of the animal.
The neck of T. rex formed a natural S-shaped curve like that of other theropods, but was short and muscular to support the massive head. The two-fingered forelimbs were very small relative to the size of the body, but heavily built. In contrast, the hindlimbs were among the longest in proportion to body size of any theropod. The tail was heavy and long, sometimes containing over forty vertebrae, in order to balance the massive head and torso. To compensate for the immense bulk of the animal, many bones throughout the skeleton were hollow. This reduced the weight of the skeleton while maintaining much of the strength of the bones.
Classification
Tyrannosaurus is the type genus of the superfamily Tyrannosauroidea, the family Tyrannosauridae, and the subfamily Tyrannosaurinae. Other members of the tyrannosaurine subfamily include the North American Daspletosaurus and the Asian Tarbosaurus, both of which have occasionally been synonymized with Tyrannosaurus. Tyrannosaurids were once commonly thought to be descendants of earlier large theropods such as megalosaurs and carnosaurs, although more recently they were reclassified with the generally smaller coelurosaurs.
In 1955, Soviet paleontologist Evgeny Maleev named a new species, Tyrannosaurus bataar, from Mongolia. By 1965, this species had been renamed Tarbosaurus bataar. Despite the renaming, many phylogenetic analyses have found Tarbosaurus bataar to be the sister taxon of Tyrannosaurus rex, and it has often been considered an Asian species of Tyrannosaurus. However, a recent redescription of the skull of Tarbosaurus bataar has shown that it was much narrower than that of Tyrannosaurus rex and that during a bite, the distribution of stress in the skull would have been very different, closer to that of Alioramus, another Asian tyrannosaur. A related cladistic analysis found that Alioramus, not Tyrannosaurus, was the sister taxon of Tarbosaurus, which, if true, would suggest that Tarbosaurus and Tyrannosaurus should remain separate.
Other tyrannosaurid fossils found in the same formations as T. rex were originally classified as separate taxa, including Aublysodon and "Albertosaurus" megagracilis. However, these fossils are now universally considered to belong to juvenile T. rex. A small but nearly complete skull from Montana, 60 cm (2 ft) long, may be an exception. This skull was originally classified as a species of Gorgosaurus ("G." lancensis) by Charles W. Gilmore in 1946, but was later referred to a new genus, Nanotyrannus. Opinions remain divided on the validity of N. lancensis. Many paleontologists consider the skull to belong to a juvenile T. rex. There are minor differences between the two species, including the higher number of teeth in N. lancensis, which lead some scientists to recommend keeping the two genera separate until further research or discoveries clarify the situation.
Manospondylus controversy
The first fossil specimen which can be attributed to Tyrannosaurus rex consists of two partial vertebrae (one of which has been lost) found by Edward Drinker Cope in 1892 and described as Manospondylus gigas. Osborn recognized the similarity between M. gigas and T. rex as early as 1917 but, due to the fragmentary nature of the Manospondylus vertebrae, he could not synonymize them conclusively.
Controversy erupted in June 2000 after more tyrannosaur bones unearthed in South Dakota by the Black Hills Institute were found at the type locality of M. gigas and judged to represent further remains of the same individual. These more recently-discovered remains clearly belong to T. rex. According to the rules of the International Commission on Zoological Nomenclature, the system that governs the scientific naming of animals, Manospondylus gigas should therefore have priority over Tyrannosaurus rex, because it was named first. However, in the Fourth Edition of the International Code of Zoological Nomenclature, which took effect on January 1 2000, Chapter 6, Article 23.9 states that "the prevailing usage must be maintained" when "the senior synonym or homonym has not been used as a valid name after 1899" and "the junior synonym or homonym has been used for a particular taxon, as its presumed valid name, in at least 25 works, published by at least 10 authors in the immediately preceding 50 years..." Tyrannosaurus rex more than qualifies as the valid name under these conditions and is considered a nomen protectum ("protected name") under the ICZN, making Manospondylus gigas a nomen oblitum ("forgotten name").
Paleobiology
As with all dinosaurs known only from the fossil record, much of Tyrannosaurus biology, including behavior, coloration, ecology, and physiology, remains unknown. However, many new specimens have been discovered in the last twenty years, which has allowed some informed speculation on growth patterns, sexual dimorphism, biomechanics, and metabolism.
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Superorder: Dinosauria
Order: Saurischia
Suborder: Theropoda
Family: Tyrannosauridae
Subfamily: Tyrannosaurinae
Genus: Tyrannosaurus
Species: T. rex
Tyrannosaurus meaning 'tyrant lizard') is a genus of tyrannosaurid theropod dinosaur. The species Tyrannosaurus rex, commonly abbreviated to T. rex, is one of the dinosaurs most often featured in popular culture around the world. It hails from what is now western North America. Some scientists consider the slightly older Tarbosaurus bataar from Asia to represent a second species of Tyrannosaurus, while others maintain Tarbosaurus as a separate genus.
Like other tyrannosaurids, Tyrannosaurus was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to the large and powerful hindlimbs, Tyrannosaurus forelimbs were small and retained only two digits. Although other theropods rivaled or exceeded T. rex in size, it was the largest known tyrannosaurid and one of the largest known land predators, measuring over 12 metres (40 feet) in length and weighing as much as an elephant.
Fossils of some T. rex have been found in North American rock formations dating to the very end of the Cretaceous Period (late Maastrichtian stage, 65 million years ago); it was among the last dinosaurs to exist prior to the Cretaceous-Tertiary extinction event. More than 30 specimens of T. rex have now been identified, some nearly complete, which has allowed significant research into many aspects of its biology, including its life history and biomechanics. The feeding habits and potential speed of T. rex remain controversial.
Description
Tyrannosaurus rex was one of the largest land carnivores of all time, about 12 to 13 meters (40 to 43.3 feet) long, and 4.5-5 m (14-16 ft) tall, when fully-grown.[1] Mass estimates have varied widely over the years, from more than 7,200 kilograms (8 short tons), to less than 4,500 kg (5 tons), with most modern estimates ranging between 5,400 and 6,800 kg (between 6 and 7.5 tons).
The largest known T. rex skulls measure up to 1.5 m (5 ft) in length. Compared to other theropods, the skull was heavily modified. The skull was extremely wide posteriorly, with a narrow snout, allowing some degree of binocular vision. Some of the bones, such as the nasals, were fused, preventing movement between them. Large fenestrae (openings) in the skull reduced weight and provided areas for muscle attachment. The bones themselves were massive, as were the serrated teeth which, rather than being bladelike, were oval in cross-section. Like other tyrannosaurids, T. rex displayed marked heterodonty, with the premaxillary teeth at the front of the upper jaw closely-packed and D-shaped in cross-section. Large bite marks found on bones of other dinosaurs indicate that these teeth could penetrate solid bone. T. rex had the greatest bite force of any animal. Worn or broken teeth are often found, but unlike those of mammals, tyrannosaurid teeth were continually replaced throughout the life of the animal.
The neck of T. rex formed a natural S-shaped curve like that of other theropods, but was short and muscular to support the massive head. The two-fingered forelimbs were very small relative to the size of the body, but heavily built. In contrast, the hindlimbs were among the longest in proportion to body size of any theropod. The tail was heavy and long, sometimes containing over forty vertebrae, in order to balance the massive head and torso. To compensate for the immense bulk of the animal, many bones throughout the skeleton were hollow. This reduced the weight of the skeleton while maintaining much of the strength of the bones.
Classification
Tyrannosaurus is the type genus of the superfamily Tyrannosauroidea, the family Tyrannosauridae, and the subfamily Tyrannosaurinae. Other members of the tyrannosaurine subfamily include the North American Daspletosaurus and the Asian Tarbosaurus, both of which have occasionally been synonymized with Tyrannosaurus. Tyrannosaurids were once commonly thought to be descendants of earlier large theropods such as megalosaurs and carnosaurs, although more recently they were reclassified with the generally smaller coelurosaurs.
In 1955, Soviet paleontologist Evgeny Maleev named a new species, Tyrannosaurus bataar, from Mongolia. By 1965, this species had been renamed Tarbosaurus bataar. Despite the renaming, many phylogenetic analyses have found Tarbosaurus bataar to be the sister taxon of Tyrannosaurus rex, and it has often been considered an Asian species of Tyrannosaurus. However, a recent redescription of the skull of Tarbosaurus bataar has shown that it was much narrower than that of Tyrannosaurus rex and that during a bite, the distribution of stress in the skull would have been very different, closer to that of Alioramus, another Asian tyrannosaur. A related cladistic analysis found that Alioramus, not Tyrannosaurus, was the sister taxon of Tarbosaurus, which, if true, would suggest that Tarbosaurus and Tyrannosaurus should remain separate.
Other tyrannosaurid fossils found in the same formations as T. rex were originally classified as separate taxa, including Aublysodon and "Albertosaurus" megagracilis. However, these fossils are now universally considered to belong to juvenile T. rex. A small but nearly complete skull from Montana, 60 cm (2 ft) long, may be an exception. This skull was originally classified as a species of Gorgosaurus ("G." lancensis) by Charles W. Gilmore in 1946, but was later referred to a new genus, Nanotyrannus. Opinions remain divided on the validity of N. lancensis. Many paleontologists consider the skull to belong to a juvenile T. rex. There are minor differences between the two species, including the higher number of teeth in N. lancensis, which lead some scientists to recommend keeping the two genera separate until further research or discoveries clarify the situation.
Manospondylus controversy
The first fossil specimen which can be attributed to Tyrannosaurus rex consists of two partial vertebrae (one of which has been lost) found by Edward Drinker Cope in 1892 and described as Manospondylus gigas. Osborn recognized the similarity between M. gigas and T. rex as early as 1917 but, due to the fragmentary nature of the Manospondylus vertebrae, he could not synonymize them conclusively.
Controversy erupted in June 2000 after more tyrannosaur bones unearthed in South Dakota by the Black Hills Institute were found at the type locality of M. gigas and judged to represent further remains of the same individual. These more recently-discovered remains clearly belong to T. rex. According to the rules of the International Commission on Zoological Nomenclature, the system that governs the scientific naming of animals, Manospondylus gigas should therefore have priority over Tyrannosaurus rex, because it was named first. However, in the Fourth Edition of the International Code of Zoological Nomenclature, which took effect on January 1 2000, Chapter 6, Article 23.9 states that "the prevailing usage must be maintained" when "the senior synonym or homonym has not been used as a valid name after 1899" and "the junior synonym or homonym has been used for a particular taxon, as its presumed valid name, in at least 25 works, published by at least 10 authors in the immediately preceding 50 years..." Tyrannosaurus rex more than qualifies as the valid name under these conditions and is considered a nomen protectum ("protected name") under the ICZN, making Manospondylus gigas a nomen oblitum ("forgotten name").
Paleobiology
As with all dinosaurs known only from the fossil record, much of Tyrannosaurus biology, including behavior, coloration, ecology, and physiology, remains unknown. However, many new specimens have been discovered in the last twenty years, which has allowed some informed speculation on growth patterns, sexual dimorphism, biomechanics, and metabolism.
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Superorder: Dinosauria
Order: Saurischia
Suborder: Theropoda
Family: Tyrannosauridae
Subfamily: Tyrannosaurinae
Genus: Tyrannosaurus
Species: T. rex
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